Known only by the gold tooth in his mouth, this elderly alchemist is a ruthless servant of Father and one of the minds behind King Bradley's making. Armed with his own Philosopher's Stone and privy to the nation's darkest secrets, he treats human lives as raw material for engineering a new kind of being.
The doctor is a short, aged man marked most obviously by a single golden tooth. He keeps his auburn hair cropped short beneath a small mustache and usually appears in a white lab coat with the sleeves pushed up past the elbows, spectacles perched on his nose. Unsettlingly, his gaze often drifts toward the corners of his eyes, giving him a cross-eyed, faintly unhinged look.
Brilliant but insufferably arrogant, the doctor thinks of himself as the father of Bradley and the other test subjects, regarding them less as people than as disposable projects on the road to manufacturing a new human being. He is utterly without conscience, ready to hurt, threaten, or kill anyone in his way, and indifferent to the fate of humanity at large. Gladly complicit in his own country's ruin, he plays the part of the classic mad scientist who fancies himself above others, perhaps even godlike for toying with life. That self-regard blinds him to the obvious: the Homunculi he serves still count him as nothing more than a useful human, and one wholly beneath them.
He first surfaces in a flashback set some forty years before the main story, personally injecting a liquefied Philosopher's Stone into the candidate who becomes King Bradley. In the present timeline he turns up at Briggs alongside General Raven, brought in to heal Solf J. Kimblee after his fight with Scar. His Stone lets him mend grave wounds much as Dr. Marcoh can, and he commands a squad of leftover Führer candidates, veteran fighters nearly as capable as Wrath himself.
When Edward Elric, Roy Mustang, Scar, and Riza Hawkeye search Central's maze of tunnels for Father's chamber, the doctor ambushes them, having correctly guessed their route. After loosing his swordsmen, he triggers a transmutation circle that, working with others planted across the Five Laboratories, drags Edward, Alphonse, and Izumi Curtis into the Gate. His men pin down the rest while he pressures Mustang to perform human transmutation, even ordering Hawkeye's throat cut to force the issue, but Mustang refuses on her own dying command. Reinforcements led by Jerso subdue him, yet his triumph is short-lived: as he praises the newly arrived Wrath, Pride stabs him from behind and spends him as a living sacrifice. When the transmutation clears, all that remains of the doctor is a grotesque lump of flesh, his gold tooth and crooked eyes the only recognizable features, a fitting irony for a man obsessed with human transmutation.

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The Gold-Toothed Doctor is an elderly alchemist who serves Father and helped engineer King Bradley, marked by a single golden tooth and a habit of treating human lives as raw material for experimentation.
The Gold-Toothed Doctor personally injected a liquefied Philosopher's Stone into the candidate who became King Bradley, acting as one of the men behind the future Fuhrer's creation roughly forty years before the main story.
The Gold-Toothed Doctor wields his own Philosopher's Stone, which lets him heal grave wounds much like Dr. Marcoh, and he commands a squad of leftover Fuhrer candidates nearly as capable as Wrath.
The Gold-Toothed Doctor is stabbed from behind by Pride and used as a living sacrifice for a forced transmutation, leaving only a grotesque lump of flesh with his gold tooth as the sole recognizable feature.
The Gold-Toothed Doctor pressures Roy Mustang to perform human transmutation, even ordering Riza Hawkeye's throat cut to force the issue, but Mustang refuses on her own dying command.
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