Cytomander

Of the four generals, Cytomander passes most easily for human. Only two details betray the bird stock in his genome: a crest rising from his head like a rooster's comb, and the peacock plumage that adorns him.

At court Cytomander plays the aristocrat, offering Lordgenome and his fellow generals courteous, formal address. Combat exposes the fault line in that poise. Facing defeat at the hands of Team Dai-Gurren, he degenerates into a cackling lunatic, his features twisting into grotesque exaggeration. The madness passes as quickly as it arrives; back within Teppelin's walls, the polished manner simply resumes.

Cytomander takes the field after Adiane falls, personally pursuing Team Dai-Gurren aboard the Dai-Gunten. The engagement mauls his flying battleship badly enough to force a withdrawal. One month later, on day six of the human assault on Teppelin, a maddened Cytomander returns for a coordinated two-front offensive alongside Guame in the Dai-Gundo. The gambit kills him: as the Dai-Gurren drives through the Dai-Gunten to sink it, it also rams the Shuzack, Cytomander's personal Gunmen, ending the general along with his flagship.

His name fuses cytosine, one of the chemical bases of DNA, with the elemental salamander, and reads aloud as See-toh-man-dra. The Shuzack takes its own name from Suzaku, the mythical Vermilion Bird. In the English dub the on-screen spelling appears as Cytomanter, even though the spoken lines keep the Cytomandra pronunciation.