Arc-Gurren Lagann is what results when Simon fuses his Gurren-Lagann with the Arc-Gurren battleship. Born in the heat of battle to save the vessel from the Anti-Spirals, it later becomes the very engine that drives the mighty Super Galaxy Gurren Lagann.
The machine borrows its name from the Arc-Gurren that Rossiu Adai christened, and it forms the moment Simon locks that battleship together with his own Gurren-Lagann. The union is staggering in scale, with estimates placing it near five thousand meters given that it stands roughly alongside the Dekabutsu, and the Final Drill sequence scales it even further to a size many times that of Gurren Lagann itself. Like Lagann in its plainest state, this form carries no sunglasses on either of its faces.
The combination is triggered to rescue Arc-Gurren when a pair of massive Mugann close in, and its first act tears an opening in the fabric of spacetime that hurls the fused enemies to oblivion. It later strains to hold the moon back from an early crash, and in the film retelling it snaps an arm off Cathedral Lazengann with a Giga Drill Break before punching its drill toward the moon's core. It cannot pierce Ashtanga's forcefield, yet it eventually transforms into the Ultra Spiral Powerplant that feeds Super Galaxy Gurren Lagann. Its named techniques include the Space-Time Shattering Burst Spinning Punch, which rips a hole in reality, the beam-firing Vortex Cannon Maximum, the film-only Arc-Gurren Lagann Giga Drill Break, and the Super Inazuma Drill Kick.
Simon and Viral share the cockpit as its principal pilots, calling out the combination together as they fuse the machines. Gimmy Adai and Darry Adai take the controls for a stretch during the effort to hold back the moon, and Artenborough Cortich releases the Vortex Cannon Maximum that scatters much of the Mugann swarm.

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Arc-Gurren Lagann is the machine formed when Simon fuses his Gurren-Lagann with Rossiu's battleship Arc-Gurren to save the vessel from an Anti-Spiral attack, and it later becomes the engine that powers Super Galaxy Gurren Lagann.
Arc-Gurren Lagann is estimated at around five thousand meters, roughly the scale of the Dekabutsu, and the Final Drill sequence depicts it as many times larger still.
Simon and Viral serve as Arc-Gurren Lagann's principal pilots, sharing the cockpit and calling out the combination together, with Gimmy and Darry Adai briefly taking the controls during the effort to hold back the moon.
Arc-Gurren Lagann's named techniques include the reality-tearing Space-Time Shattering Burst Spinning Punch, the beam-firing Vortex Cannon Maximum, and the Super Inazuma Drill Kick.
Arc-Gurren Lagann eventually transforms into the Ultra Spiral Powerplant that feeds Super Galaxy Gurren Lagann.
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