
The Ghost Princess of the Thriller Bark Pirates, Perona once commanded its Wild and Surprise Zombies. After Kuma scattered her to Kuraigana Island, she spent the timeskip there and reluctantly nursed Zoro through his training under Mihawk.
Perona is a petite young woman with large round black eyes and uneven lower lashes that read almost like drawn-on liner. Waist-length pink hair is gathered into a pair of pigtails fastened with flower hairpins colored black and white, and she crowns her head with a red piece bearing a black cross. Her early outfit pairs a white long-sleeved top patterned with three black hearts, cropped to bare her midriff, with a red mini cape, striped black-and-white tights, a mini-skirt, a heart-buckled belt, and red buckled boots, while she keeps her toes turned inward and totes a devil-shaped parasol. Two years on, the look matures into a long black strapless gown trimmed with lavender frills, red heels, and a wide floral-decorated top hat, a pink bat inked on her left arm, her hair now wound into several spiral braids.
Childish and self-centered, Perona chases her own whims, bristles at taking orders, and demands obedience from anyone around her. She adores cute things and will only employ servants she finds adorable, dreaming of a kingdom peopled by cute creatures turned into zombies that answer to her alone; she silences Kumashi because his gruff voice clashes with his sweet looks. Cockroaches terrify her so badly that she screams and weeps at the sight, and a streak of naivety lets simple tricks rattle her completely. She is fiercely sure her Devil Fruit can topple anyone, even briefly squaring up against Kuma, yet she grew softer over the timeskip, still clinging to a teddy bear and tearfully thanking Mihawk for sheltering her. Her trademark laugh, drawn from her fruit's name, rings out as horo-horo-horo.
Ranked among the Mysterious Four, Perona once commanded a host of zombies, and Hogback openly valued her presence in Moria's ranks. She is physically sturdier than she looks, hauling the oversized Kumashi about while levitating, and she picked up cooking and navigation during her years with Mihawk, even crossing from Paradise into the New World on her own. Wielding the Horo Horo no Mi, classed as Paramecia, she conjures and steers ghosts: Negative Hollows hollow out a victim's spirit and crush their will to live, Mini and larger Hollows burst with destructive force, and a ghost network lets her spy from afar. She can also project her own spirit free of her body, passing through walls untouched and resizing herself at will, though her abandoned body lies helpless and exposed while she does so.

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The wiki does not describe a romantic relationship between Perona and Zoro; it notes only that she reluctantly nursed him through his training under Mihawk on Kuraigana Island during the timeskip.
Perona began as the Ghost Princess of the antagonist Thriller Bark Pirates, but she later became an ally to Roronoa Zoro after the timeskip.
Perona is 25 years old, having debuted at age 23 in the story.
After Bartholomew Kuma scattered her away from Thriller Bark, Perona landed on Kuraigana Island, Mihawk's home, and spent the two-year timeskip sheltering there while he hosted her.
Perona ate the Horo Horo no Mi, a Paramecia fruit that lets her summon ghosts to drain a victim's will to live or explode with force, and lets her project her own spirit free of her body.
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